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Beyond Parkin, additional E3 ubiquitin ligases such as SMAD-specific E3 ubiquitin protein ligase 1 (SMURF1), glycoprotein 78 (Gp78), mitochondrial ubiquitin ligase 1 (MUL1), seven in absentia homolog 1 (SIAH1), and Ariadne RBR E3 ubiquitin protein ligase 1 (ARIH1) also contribute to mitophagy regulation through ubiquitination of mitochondrial proteins [83]